J. Cullen & V. H. Heywood:

Notes on the European Species of Paeonia (1964)

P. officinalis L.

(a) subsp. officinalis

(b) subsp. villosa (Huth) Cullen & Heywocd, stat.nov.

(c) subsp. humilis (Retz.) Cullen & Heywood, comb. et stat. nov.

(d) subsp. banatica (Rochel) Soó,

P. mascula (L.) Mill.

(a) subsp. mascula

(b) subsp. russii (Biv.) Cullen & Heywood, comb. et stat. nov.

The account of Paeonia by J. Cullen and V.H. Heywood in Flora Europaea 1: 243-244 (1964) recorded three species from mainland Greece and Crete, namely P. peregrina, P. clusii and P. mascula. Davis and Cullen in Flora of Turkey 1: 204-206 (1965) recorded from the East Aegean Greek islands P. mascula subsp. arietina from Samos and P. rhodia from Rhodes. None made use of the material in Greek herbaria, to which numerous specimens have been added since 1964. Indeed the collections now available, made in particular by Elli Stamatiadou and Dimitrios Tzanoudakis, have established the general patterns of distribution so well that further collecting is unlikely to modify these greatly (Figs 22, 23).

From the book:

Peonies of Greece

A taxonomic and historical Survey of the Genus Paeonia in Greece

William T. Stearn and Peter H. Davis